The Doctor and Rose materialise the TARDIS in a massive underground bunker near Salt Lake City, Utah in the year 2012, drawn to it by a distress signal. They find they are in the Vault, owned by the wealthy Henry van Statten who collects and houses alien artifacts within it. While Rose tours the facility with one of van Statten's technicians, Adam Mitchell, van Statten offers to take the Doctor to see the pride of his collection, the "Metaltron", contained in a special part of the Vault called the Cage. The Doctor is first shocked to find the Metaltron is actually a Dalek, all whom he thought destroyed from the Time War, but then laughs when he discovers it is weakened and chained down, unable to fight back. The Doctor attempts to destroy the Dalek but is stopped by van Statten guards and escorted back to his offices.
Meanwhile, Adam has led Rose to the Cage, and she also finds the Dalek, and takes pity on it. She touches its casing, allowing the Dalek to absorb her DNA and time energy from the radiation she has been exposed to as part of her travels in the TARDIS. The Dalek uses these to become re-energized, drawing power from all over the southwestern United States to recharge itself, and breaks its bonds; an evacuation of the Vault is initiated. After learning from the Internet that it is the last Dalek, the Dalek begins exterminating the guards in its way. The Dalek then communicates directly with the Doctor, explaining its ability to regenerate by means of DNA extrapolation when Rose touched it. However, as the last remaining Dalek and without a command structure to receive orders from, it is to all extents and purposes, useless. The Doctor then suggests the Dalek kill itself as to remove the presence of its race from the universe. However, the Dalek refuses, stating "The Daleks must survive". The Doctor becomes increasingly aggressive, demanding it destroy itself, but the Dalek ignores him, coldly stating "You would make a good Dalek" before cutting communications.
Adam tries to help Rose escape, but she is trapped inside the Vault when it is sealed by van Statten. The video feeds are disrupted moments before the Dalek encounters Rose, infuriating the Doctor. The Dalek, infused with human DNA, shows sympathy for Rose and spares her life for the moment, demanding its release from the Cage in exchange for Rose's life, which the Doctor is forced to allow. The Dalek makes its way to van Statten's office seeking to exterminate its captor, but Rose convinces she can offer the Dalek what it really seeks, its freedom.
The Dalek and Rose travel to the highest point in the Vault, and the Dalek shoots a hole in the ceiling, sending sunlight into the facility. As the Dalek opens its casing and experiences the sunlight directly, the Doctor appears with a weapon, ready to destroy it. Rose intervenes and warns the Doctor off, showing him that her DNA has caused the Dalek to mutate and no longer able to perform its objective. After asking Rose to order it to do so, the Dalek annihilates itself, as it believes anything different to a Dalek is wrong. In the episode's dénouement, van Statten is arrested and ordered to be mind wiped, and then dumped on the streets of "San Diego, Seattle, Sacramento, someplace beginning with 'S'". Meanwhile, the Vault will be filled with cement and sealed off. At the TARDIS, the Doctor ruefully observes that as the last survivor of the Time War, he "wins". He also tells Rose that he would be able to sense the presence of other Time Lords had they survived. As Adam warns them about the impending closure of the Vault, Rose invites him to join them in the TARDIS, much to his bewilderment.
Source: wikipedia
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